Wider paleogeographical distribution of bothremydid turtles in northern south america during the paleocene–eocene

Bothremydidae was one of the most diverse and widespread group of side-necked turtles (pleurodirans) during the Cretaceous and part of the Paleogene. In South America, the Paleogene record of bothremydids is restricted to Puentemys mushaisaensis from the middle–late Paleocene Cerrejón Formation of Colombia, Inaechelys pernambucensis from the Paleocene of Brazil, and Motelomama olssoni from the early Eocene of Perú. Here, we describe two shells of P. mushaisaensis and several other isolated bones conferred to this taxon from the upper Paleocene and lower Eocene Arcillolitas de Socha Formation found in the Socha Region, Boyacá Department of Colombia. U-Pb dating of detrital zircon from two levels from this formation indicates maximum depositional ages of 56.83±0.04 Ma and 57.2±0.5 Ma for the areniscas guía interval of the formation. The new occurrence of P. mushaisaensis in the Socha region, at least 500 km south from Cerrejón, indicates a wider biogeographical distribution of northern South America Paleocene herpetofauna, possibly helped by low topography and ecosystems connectivity via a faunistic corridor.