Ursus etruscus from the late Early Pleistocene of the Taurida cave (Crimean Peninsula)

The present work describes the dentognathic remains of Ursus etruscus Cuvier, 1823 from the recently discovered Taurida cave in central Crimea at the north Black Sea area. The bone-bearing layer of Taurida cave corresponds to the Psekupsian Faunal Assemblage of Eastern Europe and to the Late Villafranchian of Western Europe (ca. 1.8-1.5 Ma). Here, we describe unpublished ursid material unearthed during the excavations performed at the cave in 2020-2021, further comparing it with coeval chronologic and geographic sites around Europe. Our anatomical and biometrical analyses suggest the inclusion of the studied specimens in the hypodigm of the Early Pleistocene medium-sized species Ursus etruscus. The finds of the U. etruscus from the southern part of Eastern Europe provide a link between the western and eastern parts of the species range. Therefore, the finds from Crimea are important for understanding of the morphological diversity and evolution of U. etruscus which is the putative ancestor of both cave bears and brown bears. Furthermore, the study of these remains is also important for understanding the processes of the forming of the large mammal assemblages in the late Early Pleistocene and its relationships with the dispersal of the genus Homo.