Why does strength training improve endurance performance?

Objective The specificity of training principle holds that adaptations to exercise training closely match capacity to the specific demands of the stimulus. Improvements in endurance sport performance gained through strength training are a notable exception to this principle. While the proximate mechanisms for how strength training produces muscular adaptations beneficial to endurance sports are increasingly well understood, the ultimate causes of this phenomenon remain unexplored. Methods Using a holistic approach tying together exercise physiology and evolution, I argue that we can reconcile the apparent "endurance training specificity paradox." Results and Conclusions Competing selective pressures, inherited mammalian biology, and millennia of living in energy-scarce environments constrained our evolution as endurance athletes, but also imparted high muscular plasticity which can be exploited to improve endurance performance beyond what was useful in our evolutionary past.