LTR retrotransposons create transcribed retrocopies in metazoans 1

1 Shengjun Tan, Margarida Cardoso-Moreira, Wenwen Shi, Dan Zhang, Jiawei Huang, 2 Yanan Mao, Hangxing Jia, Yaqiong Zhang, Chunyan Chen, Yi Shao, Liang Leng, 3 Zhonghua Liu, Xun Huang, Manyuan Long, Yong E. Zhang 4 Key Laboratory of Zoological Systematics and Evolution & State Key Laboratory of 5 Integrated Management of Pest Insects and Rodents, Institute of Zoology, Chinese Academy 6 of Sciences, Beijing 100101, China. 7 Center for Integrative Genomics, University of Lausanne, 1015 Lausanne, Switzerland 8 Present address: Zentrum für Molekulare Biologie der Universität Heidelberg (ZMBH), 9 69120 Heidelberg, Germany. 10 University of Chinese Academy of Sciences, Beijing 100049, China 11 State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and 12 Developmental Biology, Chinese Academy of Sciences, Beijing 100101, China 13 Department of Ecology and Evolution, The University of Chicago, Chicago 60637, IL, USA. 14 Corresponding author. E-mail: zhangyong@ioz.ac.cn. 15 Running title: LTR retrotransposon mediated retroposition 16