Seasonal Effects of Environmental Change on Yellow-Bellied Marmot 1 Population Dynamics 2 Running title : Seasonal mechanisms of population dynamics 3

1 Department of Evolutionary Biology and Environmental Studies, University of Zurich, 7 Winterthurerstrasse 190, CH-8057 Zurich, Switzerland 8 2 Department of Animal and Plant Sciences, University of Sheffield, Western Bank, Sheffield S10 2TN, 9 UK 10 3 Ecology & Evolutionary Biology Department, The University of Kansas, Lawrence, KS 66045-7534, 11 USA 12 4 Department of Ecology and Evolutionary Biology, University of California Los Angeles, Los Angeles, 13 CA 90095, USA 14 5 The Rocky Mountain Biological Laboratory, Crested Butte, CO 81224, USA 15 6 School of Biological Sciences, University of Aberdeen, Aberdeen, AB24 2TZ, UK 16 7 Department of Wildlife Ecology, University of Florida, Gainesville, FL 32611, USA 17 18 19 *Corresponding author: Tel.: +34-671-246-338; email: maria.paniw@ieu.uzh.ch; ORCID ID: 0000-00022