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A New Primitive Therocephalian (Theromorpha) from the Middle Permian of Eastern Europe

Yu. A. Suchkova,V. K. Golubev

Paleontological Journal（2019）

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Abstract
The largest East European scylacosaurid therocephalian Julognathus crudelis gen. et sp. nov. is described based on materials from the middle Permian locality of Sundyr-1, Mari El Republic, Russia. Julognathus occupied the niche of the largest predator within the Sundyr tetrapod community, forming a trophic match with the herbivorous tapinocephalian.
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Key words
middle Permian,Severodvinian,therocephalians,Sundyr Assemblage,synecology,Mari El,Eastern Europe
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