The Onthophagus fuscus (Coleoptera: Scarabaeidae) species complex: an update and the description of a new species.

The Onthophagus fuscus Boucomont, 1932 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) species complex has a total of eight species, one of which is a new species (Onthophagus orizabensis new species) and three are elevated from the subspecies level (O. canescens Zunino & Halffter, 1988 new rank; O. mycetorum Zunino & Halffter, 1988 new rank; O. parafuscus Zunino & Halffter, 2005 new rank). Onthophagus navarretorum Delgado & Capistan, 1996 is included, while Onthophagus semiopacus has been removed from the species complex. The distribution map and key for the identification of species in this species complex are updated. Habitat preferences, ecological tolerance, and the restriction of this species complex to the mountains of the Mexican Transition Zone are discussed.