His primary areas of investigation include Biochemistry, Sphingolipid, Saccharomyces cerevisiae, Cell biology and Cell signaling. His research on Biochemistry focuses in particular on Mutant. His Sphingolipid research is multidisciplinary, incorporating perspectives in Saccharomyces and Yeast. He has researched Saccharomyces cerevisiae in several fields, including Serine C-palmitoyltransferase and SPTLC1. His Cell biology study incorporates themes from Sphingosine kinase, Molecular biology and Lipid signaling. His Cell signaling study combines topics from a wide range of disciplines, such as Mutation, Biophysics and Phosphorylation.