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He was an outstanding scientist, whose fundamental contributions to basic and clinical science transformed the understanding of the genetic basis of cancer. Dr. Nowell, along with his research partner, the late David Hungerford, discovered the Philadelphia chromosome in 1960. This finding, an abnormally small chromosome in the cancerous white blood cells of patients with chronic myeloid leukemia (CML), was considered a watershed moment in cancer research, demonstrating the genetic basis for cancer, which ran counter to the prevailing thought at the time. The Philadelphia chromosome, along with Nowell's model of the clinical evolution of cancer, laid the groundwork for targeted therapies for oncogene-addicted cancers.
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