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His research added much to the understanding of how fungi use hydrogen peroxide and hydroxyl radicals, both derived from oxygen, to attack lignin, hemicellulose and cellulose in wood. Typically Paul was frugal in his requests for apparatus and space, but over the next 30 years, Paul published over 75 papers, cited over 3,000 times and are still being cited some 50 times a year, and supervised 9 research students, a distinguished scientific record. Not only was Paul’s own research very sound, but he always found time to have fruitful discussions with his research students and colleagues, both in Biochemistry and in Biology, His extensive knowledge of redox systems was often to prove vital in the design of critical experiments and the interpretation of their results. His Head of Department, Professor Brian Chappell appreciated the important role played by Paul in the intellectual life of the Department and memorably described Paul as “a scholar in the true sense of the word.”
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