His primary areas of investigation include Cell biology, Synaptonemal complex, Genetics, Actin and Biophysics. His work deals with themes such as Spindle apparatus and Centrosome cycle, which intersect with Cell biology. His work on Spindle pole body as part of his general Spindle apparatus study is frequently connected to Immunoelectron microscopy, thereby bridging the divide between different branches of science.

His studies in Synaptonemal complex integrate themes in fields like Synapsis, Chromosomal crossover and Prophase. In his research, Secretion, Host cell cytosol, Cell membrane, Vacuole and Cytoplasm is intimately related to Microbiology, which falls under the overarching field of Actin. His Biophysics study integrates concerns from other disciplines, such as High pressure freezing and Microscopy.